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Dr. Bronson P. Bullock

Professor, Forest Biometrics & Quantitative Timber Management
Director, Plantation Management Research Cooperative (PMRC)
  • Ph.D., Forestry – Biometrics, February 2002  Virginia Polytechnic Institute & State University, Blacksburg, VA
  • M.S., Statistics, May 2000  Virginia Polytechnic Institute & State University, Blacksburg, VA
  • M.S., Forestry – Biometrics, December 1998  Virginia Polytechnic Institute & State University, Blacksburg, VA
  • B.S., Natural Resources Management – Forestry, May 1996  Rutgers, The State University of New Jersey – Cook College, New Brunswick, NJ
Selected Publications:



  • Burkhart, H. E., Avery, T. E., & Bullock, B. P. (2019). Forest Measurements, Sixth Edition (6th ed.). Waveland Press.  434 p.

Original Research Articles – Peer-reviewed  

  • Peay, W., Bullock, B., & Montes, C. (2023). A maximum entropy approach to defining geographic bounds on growth and yield model usage. Frontiers in Forests and Global Change, 15 pages. doi:10.3389/ffgc.2023.1215713
  • Zhao, D., Bullock, B. P., & Wang, M. (2023). Long-term dynamics of aboveground carbon stocks in managed loblolly pine plantations in the southeast United States. Forest Ecology and Management, 546, 121384. doi:10.1016/j.foreco.2023.121384
  • Young, J. B., Bullock, B. P., & Montes, C. R. (2023). Assessing Mid-rotation Loblolly Pine and Competing Vegetation Responses to Post-thin Fertilization and Herbicide Application in the Southeastern United States. Journal of Forestry. doi:10.1093/jofore/fvad035
  • Shalizi, M. N., Walker, T. D., Heine, A. J., Payn, K. G., Isik, F., Bullock, B. P., & McKeand, S. E. (2023). Performance Based on Measurements from Individual-Tree Progeny Tests Strongly Predicts Early Stand Yield in Loblolly Pine. Forest Science, 12 pages. doi:10.1093/forsci/fxad002
  • Ramirez, L., Montes, C. R., & Bullock, B. P. (2023). Modeling slash pine mortality rates incorporating responses to silvicultural treatments and fusiform rust infection rates. Forest Ecology and Management, 532, 9 pages. doi:10.1016/j.foreco.2023.120832
  • Zhao, D., Bullock, B. P., Montes, C. R., & Wang, M. (2022). Production, tree size inequality and growth dominance in loblolly pine plantations under different silvicultural management regimes. FOREST ECOLOGY AND MANAGEMENT526, 13 pages. doi:10.1016/j.foreco.2022.120594
  • Kinane, S. M., Montes, C. R., Zapata, M., Bullock, B. P., Cook, R. L., & Mishra, D. R. (2022). Influence of environmental variables on leaf area index in loblolly pine plantations. FOREST ECOLOGY AND MANAGEMENT523, 17 pages. doi:10.1016/j.foreco.2022.120445
  • Ramirez, L., Montes, C. R., & Bullock, B. P. (2022). Long-term term effect of bedding and vegetation control on dominant height of slash pine plantations in the southeastern United States. FOREST ECOLOGY AND MANAGEMENT522, 11 pages. doi:10.1016/j.foreco.2022.120479
  • Wang, M., Montes, C., Bullock, B., & Zhao, D. (2022). An Inverse Growth Curve Representation of the Clutter-Jones Stand Survival Model. Forest Science68 (3), 239-245. doi:10.1093/forsci/fxac017
  • Peay, W. S., Bullock, B. P., & Montes, C. R. (2022). Growth and yield model comparisons for mid-rotation loblolly pine (Pinus taeda L.) plantations in the southeastern US. FORESTRY95(5), 616-633. doi:10.1093/forestry/cpac014
  • Restrepo, H., Bettinger, P., & Bullock, B. P. (2022). A theoretical minimal solution for heuristics: The case of the spatial harvest timber problem. COMPUTERS & OPERATIONS RESEARCH143, 8 pages. doi:10.1016/j.cor.2022.105792
  • Restrepo, H. I., Montes, C. R., Bullock, B. P., & Mei, B. (2022). The effect of climate variability factors on potential net primary productivity uncertainty: An analysis with a stochastic spatial 3-PG model. AGRICULTURAL AND FOREST METEOROLOGY315, 16 pages. doi:10.1016/j.agrformet.2022.108812
  • Zhao, D., Montes, C. R., Bullock, B. P., Wang, M., Greene, W. D., & Borders, B. (2022). Effects of intensive fertilization, complete competition control and site quality on aboveground net primary production (ANPP) dynamics of loblolly pine plantations. FOREST ECOLOGY AND MANAGEMENT506, 10 pages. doi:10.1016/j.foreco.2021.119986
  • Raut, S., Dahlen, J., Bullock, B., Montes, C., & Dickens, D. (2022). Models to predict whole-disk specific gravity and moisture content in planted longleaf pine from cutover and old field sites. CANADIAN JOURNAL OF FOREST RESEARCH52(1), 1-11. doi:10.1139/cjfr-2021-0092
  • Kinane, S. M., Montes, C. R., & Bullock, B. P. (2021). A non-parametric framework to estimate fertilization response in loblolly pine plantations using environmental covariates. FORESTRY94(3), 417-426. doi:10.1093/foresj/cpaa044
  • Zapata-Cuartas, M., Bullock, B. P., Montes, C. R., & Kane, M. B. (2021). A Dynamic Stand Growth Model System for Loblolly Pine Responding to Mid-Rotation Treatments. FORESTS12(5), 29 pages. doi:10.3390/f12050556
  • Koirala, A., Montes, C., Bullock, B., & Wagle, B. (2021). Developing taper equations for teak (Tectona grands L.f.) trees of central lowland Nepal. Trees, Forests and People5. doi:10.1016/j.tfp.2021.100103
  • Zapata-Cuartas, M., Bullock, B. P., & Montes, C. R. (2021). A Taper Equation for Loblolly Pine Using Penalized Spline Regression. FOREST SCIENCE67(1), 1-13. doi:10.1093/forsci/fxaa037
  • Koirala, A., Montes, C. R., & Bullock, B. P. (2021). Modeling dominant height using stand and water balance variables for loblolly pine in the Western Gulf, US. FOREST ECOLOGY AND MANAGEMENT479, 10 pages. doi:10.1016/j.foreco.2020.118610
  • Walker, T. D., Bullock, B. P., Smith, B. C., & McKeand, S. E. (2020). Modeling Self-Thinning Patterns in Loblolly Pine with Provenance and Family Effects. FOREST SCIENCE66(6), 712-725. doi:10.1093/forsci/fxaa030
  • Restrepo, H., Mei, B., & Bullock, B. P. (2020). Long-Term Timber Contracts in the Southeastern United States: Updating the Primer Valuation Framework. FOREST SCIENCE66(6), 653-665. doi:10.1093/forsci/fxaa020
  • Zhao, D., Bullock, B. P., Montes, C. R., & Wang, M. (2020). Rethinking maximum stand basal area and maximum SDI from the aspect of stand dynamics. FOREST ECOLOGY AND MANAGEMENT475, 10 pages. doi:10.1016/j.foreco.2020.118462
  • Zhao, D., Bullock, B. P., Montes, C. R., Wang, M., Westfall, J., & Coulston, J. W. (2020). Long-term dynamics of loblolly pine crown structure and aboveground net primary production as affected by site quality, planting density and cultural intensity. FOREST ECOLOGY AND MANAGEMENT472, 14 pages. doi:10.1016/j.foreco.2020.118259
  • Orrego, S., Montes, C., Restrepo, H., Bullock, B. P., & Zapata, M. (2020). Modeling height growth for teak plantations in Colombia using the reducible stochastic differential equation approach. JOURNAL OF FORESTRY RESEARCH32(3), 1035-1045. doi:10.1007/s11676-020-01174-y
  • Wang, M., Bullock, B. P., Montes, C., & Zhao, D. (2020). An Empirical Examination of Dominant Height Projection Accuracy Using Difference Equation Models. FOREST SCIENCE66(3), 267-274. doi:10.1093/forsci/fxz079
  • Hall, K. B., Stape, J. L., Bullock, B. P., Frederick, D., Wright, J., Scolforo, H. F., & Cook, R. (2020). A Growth and Yield Model for Eucalyptus benthamii in the Southeastern United States. FOREST SCIENCE66(1), 25-37. doi:10.1093/forsci/fxz061
  • Restrepo, H., Montes, C., Orrego, S., Salazar, J., Bullock, B., & Montes, C. (2019). Using Biophysical Variables and Stand Density to Estimate Growth and Yield of Pinus spatula: A Case Study in Antioquia, Colombia. Journal of Forestry Research. 9:195-213.
  • Zhao, D., Bullock, B. P., Montes, C. R., Wang, M., Greene, D., & Sutter, L. (2019). Loblolly pine outperforms slash pine in the southeastern United States - A long-term experimental comparison study. Forest Ecology and Management, 450, 15 pages.
  • Gallagher, D. A., Bullock, B. P., Montes, C. R., & Kane, M. B. (2019). Whole Stand Volume and Green Weight Equations for Loblolly Pine in the Western Gulf Region of the United States through Age 15. Forest Science, 65(4), 420-428.
  • Gallagher, D. A., Montes, C. R., Bullock, B. P., & Kane, M. B. (2019). Two-Step Regression Process for Whole Stand Loblolly Pine Survival Projection and Quantifying Uncertainty. Forest Science, 65(3), 265-276.
  • Denham, S. O., Coyle, D. R., Oishi, A. C., Bullock, B. P., Heliovaara, K., & Novick, K. A. (2019). Tree resin flow dynamics during an experimentally induced attack by Ips avulsus, I. calligraphus, and I. grandicollis. Canadian Journal of Forest Research, 49(1), 53-63. doi:10.1139/cjfr-2018-0024
  • Zhao, D., Westfall, J., Coulston, J. W., Lynch, T. B., Bullock, B., & Montes, C. (2019). Additive Biomass Equations for Slash Pine Trees: Comparing Three Modeling Approaches. Canadian Journal of Forest Research, 49, 27-40.
  • Restrepo, H., Bullock, B., & Montes, C. (2019). Growth and yield drivers of loblolly pine in the southeastern U.S.: A meta-analysis. Forest Ecology and Management, 435, 205-218.
  • Green, P. C., Bullock, B. P., & Kane, M. B. (2018). Culture and Density Effects on Tree Quality in Midrotation Non-Thinned Loblolly Pine Plantations. Forests, 9(2), 16 pages.
  • Scolforo, H.F., J.R.S. Scolforo, M.R. Raimundo, J.M. Mello, A.P.B. Batista and B.P. Bullock.  2015.  Hypsometric approaches to Eucalyptus experiments in Brazil.  African Journal of Agricultural Research. 10(45):4176-4184.
  • Kim, T. J., Bullock, B. P., & Stape, J. L. (2015). Effects of silvicultural treatments on temporal variations of spatial autocorrelation in Eucalyptus plantations in Brazil. Forest Ecology and Management. 358, 90-97.
  • Kim, T. J., Bullock, B. P., & McKeand, S. E. (2015). Spatial Autocorrelation among Different Levels of Genetic Homogeneity and Spacings in Loblolly Pine. Forest Science, 61(3), 438-444.
  • Kim, T. J., Bullock, B., & Wijaya, A. (2016). Spatial distribution of above-ground biomass in Labanan Concession Forest in East Kalimantan, Indonesia. Mathematical and Computational Forestry & Natural-Resource Sciences, 8(2),27-39.
  • Egbäck, S., Bullock, B. P., Isik, F., & McKeand, S. E. (2015). Height-Diameter Relationships for Different Genetic Planting Stock of Lob lolly Pine at Age 6. Forest Science, 61(3), 424-428.
  • Wood, E. R., Bullock, B. P., Isik, F., & McKeand, S. E. (2015). Variation in Stem Taper and Growth Traits in a Clonal Trial of Loblolly Pine. Forest Science, 61(1), 76-82.
  • Smith, B. C., Bullock, B. P., Isik, F., & McKeand, S. E. (2014). Modeling genetic effects on growth of diverse provenances and families of loblolly pine across optimum and deficient nutrient regimes. Canadian Journal of Forest Research, 44(11), 1453-1461. 
  • Green, E. J., & Bullock, B. (2014). Assessing Sampling and Model Error in Standing Green Weight Estimation for a Loblolly Pine Plantation. Forest Science, 60(4), 623-627. 
Research Areas:
Research Interests:

My research interests include more traditional biometry topics, such as volume, taper, and weight equations derived for various species and regions; statistical sampling and inventory issues; and the impacts of various silvicultural treatments on stand development. Further, I have more wide ranging research interests, e.g., quantitative issues relating to the growth and yield of forest stands with an emphasis on the spatial relationships between individual trees in a stand; using Bayesian statistical methods to look at spatial matrix correlation structures (with respect to a forest stand); evaluating the differences between different levels of genetic homogeneity at the individual tree and stand-level; and modeling diameter distributions for a range forest stand conditions. I try to be very flexible on the research theses my graduate students work on and I strive to make sure that their research thesis helps to achieve their professional goals.


Professor, July 2017 – present, University of Georgia, Warnell School of Forestry and Natural Resources, Athens, GA

Director, July 2023 – present, Plantation Management Research Cooperative (PMRC)

Co-Director, July 2017 – June 2023, Plantation Management Research Cooperative (PMRC)

Associate Professor, August 2014 – June 2017, University of Georgia, Warnell School of Forestry and Natural Resources, Athens, GA

Associate Professor, July 2008 – August 2014, North Carolina State University, College of Natural Resources, Department of Forestry and Environmental Resources, Raleigh, NC

Assistant Professor, March 2002 – June 2008, NC State University, College of Natural Resources, Department of Forestry and Environmental Resources, Raleigh, NC

Research Assistant, August 1996 – February 2002, Loblolly Pine Growth & Yield Research Cooperative, Virginia Polytechnic Institute & State University, Blacksburg, VA

Assistant Biometrician, May 1996 – August 1996, Westvaco Internship, Forest Science Laboratory, Biometrics Section, Summerville, SC

Nature Department Head, 1992-1995 (summers), YMCA Camp Ockanickon, Medford, NJ

Articles Featuring Dr. Bronson P. Bullock

Note: This story is part of a special issue of The Warnell Log focused on carbon

If you want to calculate the amount of carbon stored in a tree or in forests, Warnell researcher Dehai Zhao has you covered.

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